The only genus in this clade at this time is Carolus. The molecular sequences currently available (C. C. Davis & W. R. Anderson, unpublished data) place it with little support in an unresolved clade comprising mostly other genera with lateral-winged samaras.

Genus: Carolus

Distribution: New World tropics from southern Mexico to the Lesser Antilles and South America as far south as Bolivia, Paraguay, and southern Brazil. [map]